In addition, 5 leading SNPs for TL co-localized with MPN and mapped to 4 genes (TERT, NFE2, PARP1, and ATM), 2 of which have not reached genome-wide significance (P < 5 × 10−8) in prior MPN analyses (NFE2 and PARP1) (Supplementary Data 25). Here, PARP1 is linked to myeloproliferative disorder.