GBP2 and neoplasm: Among these lncRNA-mRNA pairs, the locations of nine pairs are within 1 kb, including MSTRG.5075.2-COL4A2, MSTRG.22632.1-GBP2, MSTRG.11478.1- KIR3DL2 etc. COL4A2 is coding for the alpha-2 chain of type IV collagen, an inhibitor of angiogenesis and tumor growth [46, 47].