In humans, the 1p32.3 chromosomal region harbors multiple genes (S1 Table), including autism candidate genes; Glis Family Zinc Finger 1 (GLIS1) [25,26], Carnitine Palmitoyl Transferase 2 (CPT2) [27], Sterol Carrier Protein 2 (SCP2) [27], Ubiquitin Specific Peptidase 24 (USP24) [28], SSBP3 and SSBP3 antisense (SSBP3-AS1) [29] (https://omim.org/entry/607390) (Fig 1A). Here, GLIS1 is linked to autism.