DNAJC21 and benign prostatic hyperplasia: Furthermore, through expression analysis (Fig. 4a), EMSA assay (Fig. 4c) and Dual-LUC assay (Figs. 4d, e), we proved that GS3 was directly regulated by OsmiR159–OsGAMYBL2 at the transcriptional level, OsGAMYBL2 might function as a transcriptional factor that repress the function of GS3 in BPH resistance.