We performed colocalization analysis with the eQTL data; variation in TRIM4 gene expression across 11 cancer-related tissues was potentially responsible for the GWAS locus (SS P-value: 1.4 × 10−5−5.4 × 10−5, suggested threshold after multiple testing correction <0.002; Supplementary Fig. 6c). The gene discussed is TRIM4; the disease is cancer.