In another study of the Arabidopsis transcriptome response to iron deficiency, a gene co-expression analysis revealed a subnetwork containing RACK1A and RACK1B and the four major regulators of Fe homeostasis, including BTS, PYE, bHLH39, and bHLH101 genes as well as several central players in Fe homeostasis, such as OPT3, FRO3, NAS4, IRT1, and MTPA2. Here, CLN3 is linked to Iron deficiency anemia.