HLA-DRB1 and rheumatoid arthritis: Several HLA genes have been linked to a predisposition to RA-related lung diseases, such as HLA-DQB1*03:01, HLA-DRB1*15, and HLA-DRB1*16, as well as specific mutations in other genes, such as SFTPC, RTEL1, and TERT [94,95].