Four mQTL tagging the CpGs cg08857797 (VPS25), cg00144180 (HDAC4), cg16765088 (SYNM) and cg25536676 (DHCR24) were suggestively associated with type 2 diabetes, with an unadjusted GWAS p<0.05 (ESM Table 5). Here, HDAC4 is linked to type 2 diabetes mellitus.