BSG and hepatocellular carcinoma: β1,6-GlcNAc-branched glycan of CD147/basigin by N-acetylglucosaminyltransferase V (GnT-V) upregulated matrix metalloproteinases (MMPs) expression (e.g., MMP-1, MMP-2, and MMP-9) and enhanced their associations with integrin β1, thereby contributing to HCC metastasis [41].