We also identified three novel genes, HLA-S, ZSWIM1, and GRAP2. As a pseudogene located at the major histocompatibility complex (MHC) region, HLA-S was reported to be associated with a broad range of conditions including schizophrenia, gastric, and hepatocellular cancers. The gene discussed is GRAP2; the disease is hepatocellular carcinoma.