STR “chr1:2609035” in the intron of MMEL1 was tagged by multiple SNPs associated with sclerosis, autoimmune disease, and rheumatoid arthritis; STR “chr15:77705175” in the intron of LINGO1 was in high LD with SNPs related to neuroticism measurement, unipolar depression, and mood disorder; STR “chr10:6586116” in the intron of PRKCQ-AS1 was tagged by SNPs related to multiple traits such as asthma, eczema, and allergic rhinitis (Supplementary Data 6). Here, MMEL1 is linked to rheumatoid arthritis.