Partial cytoplasmic localization is supported by the findings that Lnc473 is a target of the microRNA miR-34a (22), which is a psychiatric disease-associated microRNA that in neurons regulates synaptic maturation (23), and functions as a thermogenic regulator in adipocytes, where it localizes to the mitochondrial–lipid droplet interphase (20). Here, PDE10A is linked to psychiatric disorder.