RUNX1 and chronic myelogenous leukemia, BCR-ABL1 positive: In addition, the BCR–ABL, Tax-1, RUNX1, and NUP98–HOXA9 (NA9) fusion proteins associated with chronic granulocytic leukemia (CML) and acute myeloid or lymphoid leukemia (AML/ALL) have also been found to cause leukemia-like traits in Drosophila lymph glands [4,5,6,7].