In particular, FUSION was used with summary statistics of the PGC2 GWAS of BD (20) to discover that genomic loci associated with BD contain fetal brain cis-regulatory elements for 22 genes, of which 12 are coding: CDK10, CHMP1A, DDHD2, GLT8D1, GNL3, GOLGA6L4, GPN1, LRRC57, NMB, PACS1, SEMA3G, and XPNPEP3 (41). Here, GPN1 is linked to Behcet disease.