Lastly, the region on BTA 18 was reported as a candidate region for tuberculosis resistance (Ring et al., 2019), milk (Cole et al., 2011; Ibeagha-Awemu et al., 2016; Benedet et al., 2019), and fertility traits (Cole et al., 2011; Gaddis et al., 2016) including spermatogenesis-associated proteins 33 and 2L (SPATA33 and SPATA2L) in the extended regions of 250 kb. Here, SPATA33 is linked to tuberculosis.