Total tumor fucosylated HLA-DRB1 exhibited weak or no association with tumoral CD4+ T cells (Fig. 7d, top and bottom), although the association was modestly increased when restricted to CD4+ T cells localized at the periphery of the tumors (Extended Data Fig. 6b,c; absolute CD4+ T numbers in Supplementary Table 3). Here, HLA-DRB1 is linked to neoplasm.