PRC1 and infection: There are three key points of this study (i) AUR1 is involved in microcolony establishment and infection thread progression in root nodule symbiosis; (ii) the TPXL-AUR1-MAP65 module regulates microtubule functions for the formation of infection thread; and (iii) Myeloblastosis (MYB) family protein, MYB3R1 acts upstream of the TPXL-AUR1-MAP65 module to promote rhizobial infection and activate expression of AUR1 (Fig. 1).