TNFRSF9 and Cachexia: Similarly, the expression levels of the FFA β-oxidation gene carnitine palmitoyltransferase (Cpt1) and browning of WAT markers, including Cd137, Zic-1 and UCP-1, were increased compared with rats without cachexia in the control group (P < 0.01; Fig. 4C-E,I) [23].