GLUD1 and neoplasm: Expressions of four transcripts including L-type amino acid transporter 1 (SLC7A5), the high-affinity L-glutamine transporter (SLC1A5), glutaminase (GLS), and glutamate dehydrogenase 1 (GLUD1) were all significantly up regulated in T3 as compared with T1 and T2 tumours (Figure 4B).