Evidence for shared susceptibility was further provided at nine loci including 2p21 (THADA), 3p14.3 (ERC2), 3p13 FOXP1, 4q34.1 (HAND-AS1), 5p15.33 (CEP72), 7p15.2 (LOC646588), 7q33 (CALD1) and 9q22.31 (BARX1) of which eight were previously discovered on individual hernia GWAS analyses. Here, CALD1 is linked to Hernia.