The previously unknown loci were annotated to the pigmentation, cardiometabolic, cancer development/progression and immune-regulatory pathways (Figs. 5,  6), whilst others are known loci for cutaneous melanoma susceptibility (ATM, and SOX6 for BCC, and GPR98, and DSTYK for both BCC and SCC). Here, DSTYK is linked to skin basal cell carcinoma.