There are also two more paper from the UK Biobank published during revision of the current paper including the association of eosinophil counts with ALOX15, CSF2RB, IL17RA, IL33, JAK2, S1PR4, SH2B3, NPAT and RMI118 as well as an abandoned preprint19 from the Scandinavian Asthma Genetic Study that includes also UK Biobank participants highlighting protein-truncating variants in FLG and IL33. Here, ALOX15 is linked to asthma.