BMAL1 and hepatocellular carcinoma: Cellular staining with the lipophilic dye BODIPY 493/503 further showed that BMAL1 silencing elevated the intracellular levels of neutral lipids in SNU-368 and SNU-739, whereas BMAL1 overexpression diminished the intracellular levels of neutral lipids in HCC cells (Figure 5B).