HNF4A and hepatocellular carcinoma: However, we did not find any differences in the mRNA expression of other markers related to HCC characterisation (Saa1, Fabp1), progenitor cells (Hnf4a, Krt19), Wnt pathway (Glul, Lgr5, Oat), metabolism (Ark1b10, Gpx2), and differentiated markers (Arg1, Ctlc, Hsp1, Yap1) in Myc-R26Met versus Alb-R26Met tumours (Figs. 3H, S2).