BECN1 and fetal growth restriction: The measurement of autophagy-associated proteins in the colon showed that the protein levels of Beclin1 and p62 and the ratio of LC3 II/I were higher in the IUGR piglets than in the NBW piglets (P < 0.05; Fig. 3c), indicating that IUGR may disturb the process of autophagosomal degradation in the colon.