PDE4A and COVID-19: The genetic association in signal-A, which was tagged by rs34536443/rs74956615, also colocalizes with the cis eQTL signal for PDE4A in artery tibial in GTEx v8 data (Fig 4A, PPH4 > 0.99 for colocalisation between the two traits and with the eQTL signal) where the severe COVID-19 risk allele, that is protective for SLE, is associated with reduced expression (Table G in S1 Text).