NTS and hydrops fetalis: Among them, 5 mQTL were concurrently seen in both HF and NN, including qGW2.3 in HMPs, qGW4.2, and qGW5 in BC1F1s, while qGW6.1 and qGW9.2 were found in BC1F1s (HF), and BC1F1s and HMPs in NN with different gene actions.