Soon after, in a study of the GRMD dog model for DMD, we identified the dysregulation of cardiac-enriched miRNA, including miR-208a/b and miR-499, and of many miRNAs originated from Dlk1-Dio3 locus (Jeanson-Leh et al, 2014). Here, DLK1 is linked to Duchenne muscular dystrophy.