MMP9 and lung cancer: For example, in bladder tissues, uc.8+ traps miR-596 and lowers the availability of this miRNA to bind MMP9 mRNA (42), while, in lung cancer cells, uc.339 acts as a decoy RNA for 3 miRNAs, including miR-339-3p, miR-663-3p, and miR-95-5p (21).