MMP9 and glioblastoma: On glioblastoma cell lines LN18 and U87, Song et al. indicated that resveratrol reduced epithelial to mesenchymal transition (EMT), expression of β-catenin, and decreased the expression of stem cell markers (Twist, Snail, MMP-2, MMP-9, Slug, and Smad) [45] (Table 1).