GYPB and hydrops fetalis: For example, M-QTLs (GYP1q24, TGNP1q24 and FGNP1q23) containing OsSRLK showed OD effect on GYP but with additive effects on TGNP and FGNP in HF, while TGNP1q24 also exhibited OD effect on TGNP in DY.