Moreover, though Dynein Axonemal Light Chain 4 (DNAL4), on which a frameshift variant was found in teratozoospermic men, has not been recorded to be involved in teratozoospermia in humans, researchers observed that a mutation in this gene in boar affected sperm motility and caused midpiece abnormalities [108]. This evidence concerns the gene DNAL4 and Abnormal sperm morphology.