When considering only the COVID-19-associated hospitalization (B2) phenotype, we identified 88 gene-trait-tissue associations corresponding to 26 unique gene-trait associations (AFF3, CCR1, CCR2, CCR3, CCR5, CDCP1, CRHR1, CXCR6, DPP9, IFNAR2, IL10RB, IL10RB-AS1, KANSL1-AS1, KCNN3, LINC02210, LRRC37A2, LRRC37A4P, MAPT, OAS1, OAS3, PDE4A, PIGK, PLEKHM1, PSMD2, THBS3, ZNF778; FDR-adjusted p ≤ 0.05 while only considering B2; Supplementary Data 1–7 for all results split by COVID-19 phenotype) across 11 genomic regions (Fig. 2a for protein-coding genes). Here, ZNF778 is linked to COVID-19.