USP10 and hepatocellular carcinoma: S100A9 promotes binding between ubiquitin specific peptidase 10 (USP10) and phosphoglycerate mutase family member 5 (PGAM5), leading to the deubiquitination and stabilization of PGAM5, which increases mitochondrial ROS levels by promoting mitochondrial fission and subsequently inducing the growth and metastasis of HCC.