LGR5 and papilloma: Based upon the studies cited above, which provided evidence for CRPV infecting the bulge region in rabbits and HPV oncogenes giving rise to lesions consisting of Lgr5 progeny, we predicted that squamous dysplastic lesions/papillomas caused by MmuPV1 would preferentially arise from Lgr5+ epithelial progenitor cells, but this is not what we found.