Examining the expression of individual genes along 12p revealed the bona fide oncogene CCND2 to be universally overexpressed in all invasive tumour histologies investigated using our conservative cut-offs (≥2 log2 fold-change and <0.01 adjusted p-value), as well as ATN1 and PTMS. KRAS, in contrast, was only overexpressed in embryonal carcinoma (Supplementary Data 17). Here, ATN1 is linked to neoplasm.