We identified 8 protein-coding genes associated with both GCT and 1 or more alcohol use behaviors: ACTR1B, PLEKHM1, LRRC37A2, CRHR1, ARHGAP27, WNT3, RTN1, and LRRC37A. Five of these genes, PLEKHM1, LRRC37A2, CRHR1, ARHGAP27, and LRRC37A, were oppositely associated with GCT and alcohol consumption (eTable 42 in Supplement 1). Here, LRRC37A2 is linked to granular cell tumor.