HLA-DQA1 and systemic lupus erythematosus: The lupus-associated variant in this SNP tags the lupus-associated classical alleles DRB1*03:01 and DRB1*15:01, and is associated with increased expression of HLA-DRB1, HLA-DQA1, and HLA-DQB1 in monocyte-derived dendritic cells56.