PSMA4 and chronic obstructive pulmonary disease: Signals for PSMA4 (proteasome 20 S subunit Alpha 4), WNT3 (Wnt Family Member 3), DCBLD1 (Discoidin, CUB And LCCL Domain Containing 1), LILRA3 (Leukocyte Immunoglobulin Like Receptor A3) and EGLN2 (Egl-9 Family Hypoxia Inducible Factor 2) were validated in a separate cohort providing tissue selective validation of COPD susceptibility loci [93].