For example, whereas a G2-cyst showing the initial stages of cell death reveals only a few PCNA-demarcated, highly basophilic clusters of spermatogonia (Fig. 4A), the adjacent cyst that is at a further stage of demise (Fig. 4B) shows the latter plus completely coalesced spermatogonia that now lack basophilia, and fragmentation of these clustered corpses. Here, PCNA is linked to cyst.