Three of these overlap with previous genome-wide scans of selection, namely the protein-coding gene DNAH14, implicated in brain compression and encoding axonemal dynein [38]; MUC4, implicated in biliary tract cancer [39]; and ZAN, which encodes a protein involved in sperm adhesion, previously implicated in balancing selection and positive selection in human populations [40]. Here, MUC4 is linked to biliary tract neoplasm.