In addition, LINC00324, LINC01943, and HLA-DQB1-AS1 have also been reported to be involved in the development of tumors such as hepatocellular carcinoma, osteosarcoma, and breast cancer (Wu et al., 2020; Vishnubalaji and Alajez, 2021; Wang et al., 2021). Here, HLA-DQB1 is linked to hepatocellular carcinoma.