Several diabetes risk variants increased the expression of genes in multiple tissues, including AP3S2 (rs4932265-T), CCDC92 (rs7978610-G), HLA-DQA2 (rs601945-G) and a lncRNA RP11-252K23.2 (Table S2, Fig. S1, rs3115960-G). Here, AP3S2 is linked to diabetes mellitus.