LMNB1 and lung carcinoma: Furthermore, nucleolin was also identified in the cytoplasm/membrane fractions (Lamin B1 negative) of all the lung cancer cell lines (Figure 4B), although to a lesser extent than in the MDA-MB-435S cell line (Figure 4B, H1975, 0.56 ± 0.15, p < 0.05; A549, 0.62 ± 0.40; p = 0.10; H441, 0.46 ± 0.16, p < 0.01), while it was undetectable in the non-tumorigenic cell line MCF12A (Figure 4B).