Similar to other cancer types (63, 64) in thymic tumors, miR-525-5p acts as a tumor suppressor, inhibiting cell growth and invasion, and inducing apoptosis by repressing the target gene, HSPA9. HSPA9 is upregulated in thymoma and thymic carcinoma and correlated with poor patient survival. Here, HSPA9 is linked to thymus neoplasm.