In another study published in 1998, and through the implementation of an immunofluorescence co-localization analyses, Renu Wadhwa and colleagues noticed a co-localization of mortalin (Grp75) and p53 in the murine NIH/3T3 cells (immortalized by the gene overexpression of mot-2 form of murine mortalin), A-172 glioblastoma, U-2 OS osteosarcoma and HeLa cervical carcinoma cells. Here, HSPA9 is linked to cervical carcinoma.