In addition, abnormal expression of other kinds of GTs has been described in HCC, including C1GALT1, GALNT1, GALNT2, GALNT4, MGAT4A, MGAT5, B3GALT5, B4GALT4, ST3GAL1, ST3GAL2, ST3GAL6 and ST6GAL1. Here, ST3GAL1 is linked to hepatocellular carcinoma.