MUC4 and pancreatic neoplasm: In line with this hypothesis, measurement of the membrane-associated mucins MUC1, MUC3, MUC4, MUC12, MUC13, MUC15 and MUC16 (ref. 17) showed that MUC1 and MUC13 are expressed at high levels in pancreatic tumour cells (Extended Data Fig. 3a).