KEGG enrichment results showed that HIF-1 signaling pathway, toxoplasmosis and chemokine signaling pathway were significantly enriched in the target gene of lncRNAs in gill tissue, while complement and coagulation cascades and cytokine-cytokine receptor interaction were significantly enriched in the target gene of DE lncRNAs in skin tissue (Fig. 4D, E). Here, HIF1A is linked to toxoplasmosis.