ERAP1 and Behcet disease: Following this, Kirino et al. [37] reported another GWAS evaluating 779,465 single-nucleotide polymorphisms (SNPs) in 1209 Turkish BD patients and 1278 healthy controls, resulting in the identification of novel genetic variants in STAT4 (signal transducer and activator of transcription 4) and ERAP1 (endoplasmic reticulum amino peptidase 1).